The Lambda being that large permanent firework in the sky?

[It was strange for him, actually. Seeing the night sky so clearly was still a bit of a privilege.]

Decepticons are the faction that I lead, one side of the Great Cybertronian War. [This is like an Axiom future history lesson.] This war links many universes, many Cybertrons, and many Earths in a similar conflict. [He assumes you are human, so he makes certain to include that last bit.] My inverse similarly leads the Decepticons, but he leads them in a manner akin to my enemies, the Autobots.

Hence the inversion.